HNOOH
Nonpolar / hydrophobic

Proline

Pro · P · 115.13 Da

Non-essential

Pyrrolidine ring

The only secondary amine. The ring fixes the backbone angle and removes the amide N-H, so proline breaks α-helices and β-sheets.

Mnemonic
P for Proline, the Prison. Its side chain loops back and locks up the nitrogen.

Ionization ladder
pKaGroupChangeCharge
1.99 α-carboxyl -COOH → -COO⁻ 0 → −1
10.96 α-amino -NH₃⁺ → -NH₂ +1 → 0

pI = (1.99 + 10.96) / 2 = 6.48, the midpoint of the two pKa values that bracket the neutral species.

Net charge across pH
pH 1+1
pH 40
pH 70
pH 100
pH 13−1

A group is deprotonated once the pH climbs past its pKa. Add the contributions and you have the net charge, with no memorization required.