H₂NOOHSH
Polar, uncharged

Cysteine

Cys · C · 121.16 Da

Sulfur-containing Non-essential

Thiol (-SH)

Serine with sulfur in place of oxygen. Two cysteines oxidize to a disulfide bond, the main covalent cross-link in secreted proteins.

Mnemonic
C for Cysteine, and C for the Cross-links it forms.

Ionization ladder
pKaGroupChangeCharge
1.96 α-carboxyl -COOH → -COO⁻ 0 → −1
8.18 side chain neutral → deprotonated 0 → −1
10.28 α-amino -NH₃⁺ → -NH₂ +1 → 0

pI = (1.96 + 8.18) / 2 = 5.07, the midpoint of the two pKa values that bracket the neutral species.

Net charge across pH
pH 1+1
pH 40
pH 70
pH 10−1
pH 13−2

A group is deprotonated once the pH climbs past its pKa. Add the contributions and you have the net charge, with no memorization required.